著者名,論文名,雑誌名,ISSN,出版者名,出版日付,巻,号,ページ,URL,URL(DOI) MERILA J,Genetic architecture of fitness and nonfitness traits : empirical patterns and development of ideas,Heredity,,,1999,83,,103-109,https://cir.nii.ac.jp/crid/1574231875627839616,