ALIX- and ESCRT-III–dependent sorting of tetraspanins to exosomes

  • Jorge Larios
    Department of Biochemistry, Université de Genève, Geneva, Switzerland
  • Vincent Mercier
    Department of Biochemistry, Université de Genève, Geneva, Switzerland
  • Aurélien Roux
    Department of Biochemistry, Université de Genève, Geneva, Switzerland
  • Jean Gruenberg
    Department of Biochemistry, Université de Genève, Geneva, Switzerland

書誌事項

公開日
2020-02-12
権利情報
  • https://creativecommons.org/licenses/by/4.0/
DOI
  • 10.1083/jcb.201904113
公開者
Rockefeller University Press

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説明

<jats:p>The intraluminal vesicles (ILVs) of endosomes mediate the delivery of activated signaling receptors and other proteins to lysosomes for degradation, but they also modulate intercellular communication when secreted as exosomes. The formation of ILVs requires four complexes, ESCRT-0, -I, -II, and -III, with ESCRT-0, -I, and -II presumably involved in cargo sorting and ESCRT-III in membrane deformation and fission. Here, we report that an active form of the ESCRT-associated protein ALIX efficiently recruits ESCRT-III proteins to endosomes. This recruitment occurs independently of other ESCRTs but requires lysobisphosphatidic acid (LBPA) in vivo, and can be reconstituted on supported bilayers in vitro. Our data indicate that this ALIX- and ESCRT-III–dependent pathway promotes the sorting and delivery of tetraspanins to exosomes. We conclude that ALIX provides an additional pathway of ILV formation, secondary to the canonical pathway, and that this pathway controls the targeting of exosomal proteins.</jats:p>

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