NADP Regulates the Yeast <i>GAL</i> Induction System

  • P. Rajesh Kumar
    Cold Spring Harbor Laboratory, 1 Bungtown Road, Cold Spring Harbor, NY11724, USA.
  • Yao Yu
    Cold Spring Harbor Laboratory, 1 Bungtown Road, Cold Spring Harbor, NY11724, USA.
  • Rolf Sternglanz
    Cold Spring Harbor Laboratory, 1 Bungtown Road, Cold Spring Harbor, NY11724, USA.
  • Stephen Albert Johnston
    Cold Spring Harbor Laboratory, 1 Bungtown Road, Cold Spring Harbor, NY11724, USA.
  • Leemor Joshua-Tor
    Cold Spring Harbor Laboratory, 1 Bungtown Road, Cold Spring Harbor, NY11724, USA.

書誌事項

公開日
2008-02-22
DOI
  • 10.1126/science.1151903
公開者
American Association for the Advancement of Science (AAAS)

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説明

<jats:p> Transcriptional regulation of the galactose-metabolizing genes in <jats:italic>Saccharomyces cerevisiae</jats:italic> depends on three core proteins: Gal4p, the transcriptional activator that binds to upstream activating DNA sequences (UAS <jats:sub>GAL</jats:sub> ); Gal80p, a repressor that binds to the carboxyl terminus of Gal4p and inhibits transcription; and Gal3p, a cytoplasmic transducer that, upon binding galactose and adenosine 5′-triphosphate, relieves Gal80p repression. The current model of induction relies on Gal3p sequestering Gal80p in the cytoplasm. However, the rapid induction of this system implies that there is a missing factor. Our structure of Gal80p in complex with a peptide from the carboxyl-terminal activation domain of Gal4p reveals the existence of a dinucleotide that mediates the interaction between the two. Biochemical and in vivo experiments suggests that nicotinamide adenine dinucleotide phosphate (NADP) plays a key role in the initial induction event. </jats:p>

収録刊行物

  • Science

    Science 319 (5866), 1090-1092, 2008-02-22

    American Association for the Advancement of Science (AAAS)

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