Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, United Kingdom (S.P., S.P.V., C.A.S., A.L.P., P.H., S.G.T.); John Innes Centre, Norwich Research Park, Norwich NR4 7UH, United Kingdom (R.S., N.A.-K., A.K., M.I.B., P.N.); Commonwealth Scientific and Industrial Research Organization Plant Industry, Canberra, Australian Capital Territory 2601, Australia (P.M.C.); National Institute of Agricult (At 2011-10-19)