Cardiff School of Biosciences, Cardiff CF10 3TL, United Kingdom (A.M.P., D.F.A.O., F.M.S., R.S., R.A., H.J.R.); Warwick HRI, University of Warwick, Wellesbourne, Warwick, Warwickshire CV35 9EF, United Kingdom (V.B.-W.); and School of Biological Sciences, Royal Holloway, University of London Egham, Surrey TW20 0EX, United Kingdom (A.D.S.) (At 2008-06-06)