Michigan State University-Department of Energy Plant Research Laboratory (S.R., S.Q., K.A., P.Y., K.M.-S., R.S., J.H.), Plant Biology Department (K.A., N.L., C.G.W., A.P.M.W., J.H.), and Biochemistry and Molecular Biology Department (C.G.W.), Michigan State University, East Lansing, Michigan 48824; and Department of Molecular, Cellular, and Developmental Biology, University of Michigan, Ann Arbor (At 2009-03-27)