Centre for Sustainable Pest and Disease Management, Department of Plant Pathology and Microbiology (R.M., J.M., J.L., K.E.H.-K., J.J.R.), and Department of Biomathematics and Bioinformatics (E.L.-R.), Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, United Kingdom; Laboratory of Phytopathology, Wageningen University, 6700 EE Wageningen, The Netherlands (A.K., B.P.H.J.T.) (At 2011-04-05)