Formation and Significance of Interlocked Grain in Cinnamomum camphora I : Appearance of Interlocked Grain

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  • クスノキにおける交錯木理の形成とその意義(第1報) : 交錯木理の出現状況
  • クスノキ ニ オケル コウサク モク リ ノ ケイセイ ト ソノ イギ ダイ1ポウ コウサク モク リ ノ シュツゲン ジョウキョウ

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Interlocked grain is mainly formed in tropical broadleaved trees and also formed in Lauraceae species in the temperate regions. In this study, variations of grain angle in the radial direction of Cinnmomum camphora stems showing clear annual ring boundary were investigated. The results obtained were summarized as follows: Axial elements arranged in approximately parallel to the stem axis in the region near the pith, but began to be sloped against the stem axis with increasing annual ring numbers from the pith, and then alternately repeated S-helix and Z-helix (Figs. 1 and 2) . As interlocked grain was not recognized or was shallow in the region near the pith, the annual ring and the slope direction in that formation of interlocked grain initiated were obscured . The annual ring having clear interlocked grain was over third to tenth ring from the pith. The slope of grain in the S-helix or Z-helix reached to the peak in the vicinity of annual ring boundary at intervals of several years, and began to turn to the opposite direcrion (Figs. 1 and 3) . There was no significant relation between intervals from the peak to the next peak of grain angle and the distance or the ring numbers. Variation patterns of grain angle from bark to wood near the cambial zone were obtained (Fig. 4). It was clarified that 1) interlocked grain-like-one formed in the bark, 2) the grain angle in the cambial zone was different among sample trees, and 3) it was almost the same value as in the phloem cells and xylem cells adjacent to the cambium. This means that the slope direction of cambial cells corresponds to the elongation direction of newly formed phloem and xylem cells. Therefore, the reason for the formation of interlocked grain was that the cambial cells periodically repeated S-helix and Z-helix.

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