Revision of the Palearctic Species of the Myrmecophilous Genus Pella : Coleoptera, Staphylinidae, Aleocharinae

Maruyama Munetoshi

A biosystematic study of the Palearctic species of the myrmecophilous genus Pella (Coleoptera, Staphylinidae, Aleocharinae, Lomechusini, Myrmedoniina) is presented. The adult morphology of Pella is investigated into details in comparison with several other taxa of the Lomechusini. Pella is taxonomically revised, and is arranged as a distinct genus. Myrmelia, Pellochromonia and Lepla, formerly treated as subgenera of Zyras, are synonymized with Pella. Forty-one species including 15 new species (Pella horii, P. kidaorum, P. cooterorum, P. jelineki, P. schillhammeri, P. kinomurai, P. primorskyiana, P. hlavaci, P. iberica, P. kishimotoi, P. coreana, P. plutenkoi, P. intermedia, P. masakoae, P. zhoui) are recognised in the genus, arranged into 16 species-groups and redescribed/described with bionomical and host ant information. The following junior synonymies are proposed: Zyras (Pella) micropterus (=P. jureceki), Z. (P.) almaatensis (=P. bohaci), and Z. (P.) esau (=P. cinctipennis). Seven species are excluded from Pella, newly assigned to the other genera/subgenera and redescribed. They are: Zyras (Zyras) coloratus, Z. (Z.) iridescens, Z. (Z.) quasar, Z. (Glossacantha) ceylonicus, Z. (G.) reelsi, Myrmoecia urartu, and Dvorakatheta wrasei (n. gen.). Life cycles and behaviour of the Japanese species of Pella are shortly reported based on the field and laboratory observations.

Revision of the Palearctic Species of the Myrmecophilous Genus Pella : Coleoptera, Staphylinidae, Aleocharinae

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