National Natural Science Foundation of China [31430010],[11622102],[91430217],[11421110001] ; National Basic Research Program of China (973 Program) [2014CB943500] ; National Program for Support of Top-Notch Young Professionals ; State Key Laboratory of Plant Genomics [SKLPG2011A0103] ; China Scholarship Council (CSC) exchange scholarships ; Beijing Nova Program [Z161100004916107] ; UCAS